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GEOTRUPIDAE (Scarabaeoidea)

GEOTRUPIDAE Latreille, 1802
Authors: Patrice Bouchard; Yves Bousquet; Anthony E. Davies; Miguel A. Alonso-Zarazaga; John F. Lawrence; Chris H. C. Lyal; Alfred F. Newton; Chris A. M. Reid; Michael Schmitt; S. Adam Ślipiński; Andrew B. T. Smith.

Subfamily TAUROCERASTINAE Germain, 1897

Subfamily BOLBOCERATINAE Mulsant, 1842
Tribe Athyreini Lynch Arribálzaga, 1878
Tribe Bolbelasmini Nikolajev, 1996
Tribe Bolboceratini Mulsant, 1842
Tribe Bolbochromini Nikolajev, 1970
Tribe Eubolbitini Nikolajev, 1970
Tribe Eucanthini Nikolajev, 2003
Tribe Gilletinini Nikolajev, 1990
Tribe Odonteini Shokhin, 2007
Tribe Stenaspidiini Nikolajev, 2003

Subfamily GEOTRUPINAE Latreille, 1802
Tribe Ceratotrupini Zunino, 1984
Tribe Enoplotrupini Paulian, 1945
†Tribe Cretogeotrupini Nikolajev, 1996
Tribe Geotrupini Latreille, 1802
Tribe Lethrini Oken, 1843